On the other hand, and this makes the interpretation of some data difficult, the term saxicolous is often taken (e.g. Iwats, The status and systematic position of Hypnobartlettia fontana Ochyra and the Hypnobartlettiaceae based on molecular data. ORTHOTRICHALES (BRYOPSIDA) INFERRED FROM RBCL SEQUENCE ANALYSES1 BERNARD GOFFINET,2 RANDALL J. BAYER,3 AND DALE H. VITT Department of Biological Sciences, The University of Alberta, Edmonton, Alberta, Canada T6G 2E9 The affinities as well as the circumscription of the Orthotrichaceae (Bryopsida), one of the most diverse families of mosses, have been the focus of a controversy for … Plant J. Thamnium cossyrense Bott. A phylogeny was reconstructed using neighbor-joining, maximum parsimony in PAUP*, and likelihood criteria in RAxML, and Bayesian inference in MrBayes. Later, algae, fungi and lichens were placed in a separate division Thallophyta and liverworts, mosses in division Bryophyta. Kis is a new combination of Taxithelium perplanicaule Broth. A standardized ending for superfamilies, -acanae, is proposed. This review investigat … lunata Hem. Pulchrinodus inflatus is neither part of the genus Weymouthia nor of the Meteoriaceae or Lembophyllaceae. Bryopsida Definition By far the largest class of Bryophyta (sensu stricto) (84% of families) (Goffinet et al. Distribution of this new species is restricted to three localities in southwestern and southern China. Classes de rang inférieur Takakiopsida Sphagnopsida (les sphaignes) Andreaeopsida (les andréales) Andreaeobryopsida Oedipodiopsida Polytrichopsida Tetraphidopsida Bryopsida Ne doit pas être confondu avec bryophyte . The defining features of bryophytes are that their life cycle featuring alternating haploid and diploid generations with a dominant, branched gametophyte stage. The relationships between 66 representative pleurocarpous mosses were studied using cladistic analyses. The colonization of land by plants coincided with and was most likely facilitated by the evolution of 3-dimensional (3D) growth. Thus the sporophyte of Riccia is the simplest amongst the bryophytes, with a very high proportion of fertile tissue and the sterile tissue is very small. Well-formed: Y Recommended: N NBN ID code: NHMSYS0000841361. The generic limits proposed by various authors have resulted in an overlap of family concepts, and the systematic position of genera such as Weymouthia and Pilotrichella has been unsettled since the description of these families by Brotherus (1907) and Kindberg (1897). Hydrocryphaea wardii Dixon, collected during a joint Sino-American expedition to the Gaoligongshan in 2002 and 2004, is reported new for China from Yunnan Province. Fleisch., and P. expansum (Tayl.) The fossil E. savicziae (incertae sedis) is known from Poland and U.S.S.R. Hedenas in Spain and northern Africa, Suggestions for a New Familial Classification of Pleurocarpous Mosses, Notes on the Neckeraceae (Musci). On the other hand , the members of the class Bryopsida (e.g.,Funaria,Polytrichum , Pogonatum ) have the most complex sporophyte with a very high degree of sterility. These tools are now widely used … and recognized as Amblystegiaceae s.str. Kis. Dracaena is a genus comprising of about 40-100, Citation: Peña C, Malm T (2012) VoSeq: A Voucher and DNA Sequence Web Application. ... Circulifolia are distinctive in entire to crenulate leaf apices and filiform pseudoparaphyllia (Ninh, 1984;Buck, 1998). Maps, Flickr, Encyclopedia of Life (EOL) and GBIF (by generating data-dumps that can be processed with GBIF's Integrated Publishing Toolkit). Brotherus described the Neckeraceae in 1925 and placed it into the Leucodontales, later the family has alternatively been divided into two or three separate families: the Thamnobryaceae, the Neckeraceae and the Leptodontaceae. Wagn. Five different outgroups, Acrocladium auriculatum, Trachythecium verrucosum, Trachyphyllum inflexum, Myurium hochstetteri, and Rigodium implexum, varying from supposedly closely related groups (Acrocladium) to quite distant relatives (Rigodium) were used. Fleisch., P. calcutensis Fleisch., Porotrichum fruticosum (Brid.) The geographical grouping seems to be more strongly correlated with the phylogenetic grouping than thought before. 10.The identity of Homaliodendron neckeroides (Neckeraceae, Musci), A taxonomic study of the genus Handeliobryum Broth. Scorpidium and "Hygrohypnum" ochraceum were closely related to Calliergonaceae but were not included in the family because of the lack of support. The systematic relationship of Amblystegiaceae and Donrichardsiaceae is discussed. Assessment of epiphyte nutrient capital gives a more complete and accurate idea of the aboveground vegetation pools, and supports the idea that epiphytes may play a greater role in ecosystem nutrient dynamics than has been previously considered. Phylogenetic, Phylogeny of phytochromes in ascomycetes. spectabilis Fald. The Leptodontaceae is tentatively recognized as being composed of the genera Leptodon Mohr, Caduciella Enroth, Cryptoleptodon Ren. W. R. Buck. Ji comb, nov., which has a wider distribution in Asia. These taxa have been suggested to be related to each other in an earlier overview study of the 'pleurocarpous mosses', and similar suggestions have been made in other earlier published studies. The relationships between 93 species representing the Thuidiaceae, the Amblystegiaceae, and the temperate members of the Hypnaceae are evaluated using cladistic methods. Charles Darwin pioneered modern research on phototropism by demonstratin… The genus Ochyraea should be maintained, first with regard to the gametophytic differences between O. tatrensis and Hygrohypnum, and second because Hygrohypnum, and in particular H. smithii, might be not monophyletic according to the molecular data. The gametophyte is free-living, autotrophic, and almost always composed of a leafy stem. Jaeg. Anomodon comprises two clads, one equivalent to the subgenus Pseudoanomodon Limpr., including A. longifolius, A. attenuatus, A. giraldii and A. rostratus; the other is equivalent to subgenus Anomodon and includes the species of Haplohymenium. maderense (showing almost no differences to Th. Published. melitense Bott. Ruprecht C(1), Proost S(1), Hernandez-Coronado M(2), Ortiz-Ramirez C(2), Lang D(3), Rensing SA(4), Becker JD(2), Vandepoele K(5), Mutwil M(1). maderense specimens form a polytomous branch separated from T. pandum. PLoS ONE 7(6): e39071. Ochyra. A phylogenetic tree inferred from the chloroplast (cp) DNA trnL(UAA) intron sequences of 12 pleurocarpous moss species by a maximum parsimony analysis indicates that Hypnobartlettia fontana and Cratoneuropsis relaxa belong to the Amblystegiaceae. A key to the five species and one variety of Homalia is provided. Fleisch, Homalia pennatula (Musci: Neckeraceae), a New Combination from Southeast Asia, with a Key to the Species of Homalia Author(s): Si, Análisis cladístico de la familia Anomodontaceae, The status of Thamnobryum maderense (Kindb.) Bryopsida is an extremely speciose, but fairly uniform, set of plants -- a large and distinct monophyletic assemblage. (PPTX), Botanica Orientalis Journal of Plant Science. Some suggestions for the benefits of the biodiversity and natural resource conservation in Himalaya regions will be addressed. The results of this thesis show that the Neckeraceae need re-circumscription; this includes changes in the genus composition. Classification of extant moss genera. Therefore we chose to use mainly non-coding DNA sequences from rapidly evolving DNA regions. analyses used (RAxML/NJ by PAUP/MrBayes) are shown on the respective branches. The circumscription of the Plagiotheciaceae was evaluated cladistically, using anatomical and morphological data, plus rps4 and trnL-trnF gene sequences. However, in some cases, such as the definition of the Plagiotheciaceae clade, vegetative gametophytic features proved to be more important than other characters. Their evolution by both advancement and … D Pied. The results suggest that ISSR markers may be an alternative for distinguishing sibling moss species when sequences of the most variable genomic regions traditionally used at low taxonomic level, such as ITS, do not provide the appropriate degree of polymorphism. ex Gangulee are discussed. POL II has proven to be a useful gene for resolving tick phylogeny. BRYOPSIDA Figure 1. & Lesq.) 5.5%. Phylogenetic analyses of morphological evolution in the gametophyte and sporophyte generations of the moss order Hookeriales (Bryopsida) Mol Phylogenet Evol . Porothamnium stipitatum (Mitt.) The introductory section of the book provides a discussion of the main morphological features of the hornworts, liverworts, and mosses an overview of the different regions and habitats within the Neotropics and their characteristic bryophytes, information on how to collect and process bryophytes, a list of important herbaria in tropical America, a glossary of technical terms, and a bibliography. There is a need to explore the use of nuclear protein-encoding genes because these genes direct most aspects of the phenotypic traits in the development of an organism. This lecture also explains the body structure, alternation of generation and reproduction in … The downgrade or the regressive evolution theory: seem to form morphologically heterogeneous genera. The results showed that thirty of 102 voles and two of 49 pikas were infected with E. multilocularis. M. Fleisch. The sequence of a key gene, cytochrome b, was compared among 20 Bos species and the bongo antelope, used as an outgroup. The status of Th. Based on concatenated sequences of the nad5, atp6, cox1, and nad1 genes all haplotypes were divided into two clusters. Their gametophytes are externally complex and most of them have sporophytes with greater internal differentiation of tissues. Two analyses were conducted with 37 ingroup species, representing 14 genera, and eight outgroup species. Their evolution by both advancement and reduction makes circumscription difficult, with nearly every character having exceptions. The phylogeny tree also indicated that the five species in sect. Alternatively, because the placement … According to Smith (1955) these orders are of an advanced type among the Hepaticae. The evolution of several gametophyte features previously thought to be reliable for delineating the family are also strongly correlated with habitat. It differs from all species of Porotrichum and Porothamnium in having leaves with very broad costae and short upper cells. Bryopsida. On the other hand the gametophytes of Anthocerotales, except for their embedded sex organs resemble those of thallose Hepaticae. In the analysis of the population genetic structure, clade B formed an independent phylogeographic group, but clade A was further subdivided into three groups: two covering western and eastern parts of the Korean peninsula, respectively, and the other occupying one eastern coastal islet and Japanese Tsushima Island. In the single most parsimonious tree inferred from a combined data set of both molecular markers, the genus Thamnobryum is monophyletic and separated from species of two other genera of the Thamnobryaceae (Porothamnium stipitatum, Porotrichum usagarum). Dacryophyllum falcifolium, a new North American genus and species of moss, is described and illustrated. Hypnum schmidii C.Müll. nov. differs from other species in the genus by its gonioautoicous condition and short, frequently double costae. A revised description and new illustration of H. wardii are provided. A new moss species, Homaliodendron pulchrum, from China, is delimited by its distinct morphology and biogeography. The metacestodes of E. multilocularis are fluid-filled, asexually proliferating cysts, and they are mainly found in the host's liver in the form of tumor-like growths. had very close relationships. Fleisch. It differs, however, genetically too much from Th. Results from a previous broad-scale analysis employing trnL-trnF sequence data for 168 Hypnalean and 11 Hookerialean taxa, and an analysis employing two chloroplast regions, trnL-trnF and atpB-rbcL, one nuclear region, the internal transcribed spacers of 18S-26S rDNA., plus 68 morphological characters for a reduced data set of 54 Hypnalean taxa, were used to circumscribe Amblystegiaceae. melitense (Bryopsida). A revision of the genus Echinodium recognizes six extant and one extinct species. Underlying these advances is the evolution of genes encoding specialized proteins that form novel microtubular arrays of the cytoskeleton. Image Courtesy : anbg.gov.au/bryophyte/photos-800/grrimmia-sp-exposed-rock-WA.jpg. In the present study, the number of 'islands', with trees of similar length resulting from an analysis, increased with the supposed taxonomic distance between the outgroup and the ingroup. Its distribution and morphological and ecological differences from Thamnobryum alopecurum (Hedw.) In the same publication, Bottini also described T. cossyrense var. and Glossadelphus natans (Müll. fernandesii has eight-ranked leaves and is therefore more related to Th. Bryophytes thrive in humid climates, but can be found all over the world, even in. The recent developments in laboratory techniques (namely the polymerase chain reaction and automatic sequencing) and in computer tools for the fast application of sophisticated statistics provided new, concise methods to evaluate phylogenetic relationships at any level. L' « Inventaire des bryophytes des Hauts-de-France » est un document de référence sur la flore régionale des bryophytes (mousses et hépatiques). © 2008-2020 ResearchGate GmbH. Origin and evolution of bryophytes 1. Scorpiurium sendtneri is recorded for the first time for France. Recent phylogenetic studies indicate that Homaliodendron is polyphyletic and is divided into two clades, Phylogenetic tree of the Arabidopsis and Medicago truncatula remorin protein families [45] and the three remorins from the Datisca glomerata nodule transcriptome (arrows). It also has inbuilt BLAST capabilities against all DNA sequences stored in VoSeq as well as sequences in NCBI GenBank. In the most recent comprehensive classification 28 genera were included in the Neckeraceae family. Elymus (C. nutans, E. breviaristatus, E. sinosubmuticus and E. atratus) share a close relationship. The schematic representation of the progressive evolution of gametophytes in liverworts has been depicted in Fig. Bryophytes are small, non vascular land plants, that require water for reproduction. In this study, we investigated the, FASconCAT is a user-friendly software that concatenates rapidly different kinds of sequence data into one supermatrix file. He & Snider is recorded from Madagascar the second time. The mosses (Bryopsida) also make a blindly ending evolutionary side line. A growing amount of evidence indicates that a grouping based on sporophytic characters is artificial and based on convergent evolution. The mosses are sometimes thought to have been evolved from leafy Hepaticae (i.e., Jungermanniales and Calobryales). Structures of both gametophytes and sporophytes have to be considered and in mosses these two phases have attained specialisation in different degrees in the different species. melitense Bott. Jmportant and relatively stable clades are formed by (1) the species of the Amblystegiaceae (s.l.) Berne Convention (Appendix I (continuation)), version 2 Berne Convention (Appendix I), version 2 European Topic Centre interpretation of the Habitat and Species Directive Annex 5, version 2 Common names. & Yas,; India: Thamnobryum alleghaniense (C.Müll.) The relationships of the climate change adaptation, biogeography, and conservation with the two plant genera will be further discussed in this study. FASconCAT can handle FASTA, PHYLIP and CLUSTAL formatted input files in one single run. No severe conflicts appeared between the different methods used, but the indel coding affected the support values of the inferred topologies. Bootstrap analysis with 1000 bootstrap, This research focuses on the biodiversity and the evolutionary history of the world-wide medicinal plant genus, Dracaena, and the plant genus Pleomele. The Bryopsida can be simplified into three groups: the acrocarpous (pinnate), the pleurocarpous (side-fruited), and the cladocarpous (branching) mosses. The Video lecture from Kingdom Plantae (F.Sc. In such a vast group, naturally, there are many variations so that classification is difficult. Epilithic or saxicolous bryophytes may be defined as those growing directly on the surface of rock, and in this chapter do not include aquatic species; epiphytic or corticolous species are those growing on the bark of living trees and shrubs. In 1908, Bottini published further de-scriptions and illustrations of the new taxa. tangutorum, E. purpuraristatus and E. dahuricus var. A new system of classification for recircumscribed Amblystegiaceae, including morphological delimitation of presented clades based on maximum likelihood reconstruction of ancestral character states, is proposed and appropriate nomenclatural changes made. The epiphyte communities of a Costa Rican cloud forest make up a conspicuous portion of the canopy, especially on large canopy dominants. A morphological study revealed that it clearly distinguished from Th. (EPS). Description: The Bryopsida constitute the largest class of mosses, containing 95% of all moss species. ( Thamnobryum schmidii (C.Müll.) The new species is described and illustrated and a key to the species of Homaliodendron in Eastern Asia is provided. In the Bryopsida, both columnella and spores originate from endothecium. reconstruction was employed using neighbor joining and maximum parsimony in PAUP and Bayesian inference in MrBayes 3.1. There has never been much doubt that it is the sister group of all other mosses. We also found significant branching differences in the tree topologies between wild and domestic cattle. The group of pleurocarpous mosses comprises approximately 5000 species, which corresponds to about half of all mosses. Broth,; Malaysia: Neckera warburgii Broth. In conclusion, the genetic diversity of E. multilocularis based on mt genes on a small local area is at low level but between different regions with long distance and different ecological environment each other, the genetic diversity is at relatively high level; genetic variation is higher in the nad1 gene than that in the other three mt genes. This is based on the position of the perichaetia and sporophytes. 1. The nutrient capital (in g) is: N = 3062; P = 97; K = 678; Ca = 460; Mg = 126; Na = 207. Th. Phyllodon perplanicaulis (Broth.) is synonymized with Scorpiurium circinatum (Brid.) Only a few studies have attempted to address the origin and evolution of phototropins (Briggs et al., 2001; Lariguet and Dunand, ... (Bryopsida), resulting in moss PHOT2A-C. Aulacomnium androgynum with asexual gemmae on a modified stem tip. is synonymized with Scorpiurium sendtneri (Schimp.) Hal.) Enter search terms. Jaeg. Bootstrap values >60% and posterior probabilities >0.80 for the three methods of, The high level phylogenetic relationships within Harmonia. nov.) in New Zealand and Lord Howe Island, and var. Analysis of DNA sequences from the mitochondrial COI protein-coding gene revealed 20 mtDNA-sequence-based haplotypes with a maximum divergence of. Iwats., Isopterygium, and Taxiphyllum, but resolves the rest of the Plagiotheciaceae as a monophyletic group. The ecology and distribution of the moss are discussed. are new synonyms of Phyllodon truncatulus (Müll. These leafy forms finally led to the establishment of the higher degree of internal differentiation of the tissue in Bryopsida. This initial analysis suggests a highly polyphyletic Plagiotheciaceae, with Acrocladium, Catagonium, Herzogiella, Isopterygiopsis pulchella (Hedw.) Organisms, Diversity & Evolution 8 (2008) 282– 292 Explaining the ‘anomalous’ distribution of Echinodium (Bryopsida: Echinodiaceae): Independent evolution in Macaronesia and Australasia The class Bryopsida represents the most diversified lineage, containing more than 95% of modern mosses, whereas other classes are species‐poor. The trees inferred from the POL II sequences using maximum parsimony (MP), neighbor joining (NJ), and maximum likelihood (ML) by PAUP* and MrBayes are largely concordant with the existing phylogenies. Fennici 38: 45–46. According to the latest phylogenetic studies of pleurocarpous mosses, based on molecular data, the Neckeraceae belong to the order Hypnales and share a sister group relationship with the Lembophyllaceae. Mosses (Bryophyta) are a key group occupying an important phylogenetic position in land plant (embryophyte) evolution. Shaw (2008), Morphology and classification of the Bryophyta, in Goffinet & Shaw (eds.) In contrast, Thamnobryum pandum from New Zealand differs in 7 substitutions in the trnL intron and in 3 substitutions and one indel in the ITS2. Thamnobryum cataractarum sp. In addition, in this chapter the ancestral character states of some morphological characters within the Neckeraceae are reconstructed. Porotrichum tenuinerve sp. Evolution of the Neckeraceae (Bryophyta): Resolving the backbone phylogeny. Plant evolution is marked by major advances in structural characteristics that facilitated the highly successful colonization of dry land. Adaptations for calcareous substrates evolved either in the ancestor of the entire Isopterygiopsis-Plagiothecium clade and were lost twice, or they evolved independently in Isopterygiopsis and in a clade consisting of Bardunovia, Myurella, Orthothecium, and Platydictya. Turczaninovia (C. dahurica var. Exemple de mousse Mnium hornum, Mniaceae, Mniales, Bryopsida, Bryophytes (Watermael-Boitsfort, Province de Brabant, Belgique - 08/04/1986 - Diapositive originale réalisée par Eric Walravens). Two species of Campylopus, C. fragiliformis and C. pilifer, were previously reported from Trindade Island, a Brazilian island situated in the South Atlantic Ocean around 1100 km off the coast of mainland Brazil. alopecurum than to Th. Ji gen. nov. (Neckeraceae, Bryopsida) is described from SE Asia. Light is the ultimate source of energy for almost all of life on earth, and a remarkable diversity of organisms uses photosynthesis to convert light into metabolic energy. Phylogenetic analyses of morphological evolution in the gametophyte and sporophyte generations of the moss order Hookeriales (Bryopsida). A transfer of Pilotrichella into the Lembophyllaceae is not supported. De Sloover sp. Musci. Suborders and superfamilies are used to reflect the proposed phylogeny. Anomodon is known only from the northern hemisphere: there are four species in Mexico (several of which extend to Guatemala and/or the Caribbean islands), two in the United States and Canada, on in Europe, and the remaining in south and east Asia. The molecular data does not align Pilotrichella with either the Meteoriaceae or the Lembophyllaceae sampled. The sequence (acc no JX988758) from Picomonas judraskeda as well as other newly determined sequences (acc no JX988759- JX988767) are highlighted in the tree. Plagiarism Prevention 4. Iwats., Rhizofabronia, Struckia, and Taxiphyllum placed among the outgroup taxa. 6, Berlin, Germany, D-14195 Another three analyses were performed with different sets of characters excluded to see if some kinds of characters were more important than the others for the results of the analyses. Touwia laticostata is proposed as a new genus and species of the Neckeraceae, subfamily Thamnobryoideae. and H. axyridis ab. Iwats., P. pohliaecarpum (Sull. The Marchantiales of Hapaticae is one characterized by gametophytes of external simplicity but internal complexity. Lectotypes are selected for Neckera mariei and Pinnatella microptera Fleisch. If favoured, FASconCAT dispenses output files in PHYLIP format with relaxed (unlimited signs) or restricted taxon names (up to ten signs) while sequences are printed in non-interleaved format. Sigmatella natans Müll Hal. Disclaimer 9. To read the full-text of this research, you can request a copy directly from the author. First, the rps4 (73 informative nucleotide sites) plus trnL-trnF data set (39 sites) was analyzed alone. Touw ex J.L. a new species of neckera (neckeraceae, bryopsida) from xizang, china - volume 64 issue 3 - j. enroth, m. ji Skip to main content Accessibility help We use cookies to distinguish you from other users and to provide you with a better experience on our websites. Enroth, gen. et comb. Acrocarps are generally characterized by an upright growth habit that is unbranched or only sparingly branched. De Sloover [Thamnium comorense (Hampe ex C. The results suggest that the following genera belong to the family: Acrocladium, Bardunovia, Catagonium, Herzogiella, Isopterygiopsis, Orthothecium, Myurella, Plagiothecium, Platydictya, Pseudotaxiphyllum, Rhizofabronia, and Struckia. file and the concatenated supermatrix. The three species are compared, and the habitat of the new species is described. umbrosum (E. parvulum syn. Overall, genetic divergence was similar among R. squarrosus and R. subpinnatus on the one hand and R. loreus and R. triquetrus on the other. Our study demonstrated that POL II gene sequences contain strong phylogenetic signals in ticks at the generic and higher levels. By identifying only one type of unit class for each haplome, we propose that the 5S nrDNA sequences of species within Campeiostachys may have undergone haplome‐specific concerted evolution. For example, the program provides the range information of each concatenated gene (partition) and delivers a check list of all concatenated sequences (taxa). Phyllodon lingulatus is new to Africa. The hypothesis that Hypnobartlettia fontana is a mutant of Cratoneuropsis relaxa is not supported by molecular data because of a higher similarity of the trnL(UAA) intron and the nuclear ribosomal ITS 1,2 sequences between Hypnobartlettia fontana and Amblystegium serpens than between H. fontana and Cratoneuropsis relaxa. (Echinodiaceae: Hypnobryales), The status of Thamnobryum angustifolium (Holt) Crundw, Differentiation in DNA Fingerprinting and Morphology among Species of the Pleurocarpous Moss Genus, Rhytidiadelphus (Hylocomiaceae), A New Species of Thamnobryum (Musci: Neckeraceae) from Venezuela, with a Key to the Neotropical Species of Thamnobryum, Porotrichum tenuinerve (Musci: Neckeraceae), a New Species from Honduras, with a Provisional Key to Porotrichum in Central America, Epiphyte Biomass and Nutrient Capital of a Neotropical Elfin Forest, Touwia laticostata , a remarkable new genus and species of moss from Queensland, Australia, Note de bryologie africaine XIV. Hal.) One significant contribution of this research will be in promoting molecular taxonomy to solve problems in systematics especially in cases when the classification is in debate. Discovery of the sporophyte of Himantocladium warburgii in dicates that the species belongs to Neckera Hedw. More recent ideas about the classification of the Isobryales, which are partly based on the differences between perfect and reduced peristomes, are not supported. Glossadelphus baldwinii Broth. Leptodontium, The systematic position of Gradsteinia andicola Ochyra (Donrichardsiaceae, Bryopsida) : evidence from nrDNA internal transcribed spacer sequences, Taxonomic status of Thamnium cossyrense and T. cossyrense var. Porothamnium frahmii Enroth sp. M. Fleisch. The sequences were aligned using ClustalW [90]. Organisms, Diversity & Evolution 8 (2008) 282– 292 Explaining the ‘anomalous’ distribution of Echinodium (Bryopsida: Echinodiaceae): Independent evolution in Macaronesia and Australasia T. fernandesii and T. cataractarum are retained in Thamnobryum rather than in Crassiphyllum Ochyra. It is therefore concluded that the family Donrichardsiaceae is artificial in spite of its strong morphological integrity. “the Core Pleurocarps”) form a monophylum, which consists typically of perennial mosses with creeping stems and abundant lateral branches. Symphyodon pygmaeus (Broth.) Some previous studies based on molecular data have challenged the family concept of the Neckeraceae, indicating the need for a revision of the family. According to this interpretation the bryophytes of primitive nature are found among the liverworts (Hepaticopsida-Hepaticae). DNA fingerprinting techniques including inter simple sequence repeats (ISSR) and restriction digest patterns from 18S-26S nuclear ribosomal DNA internal transcribed spacers (ITS) were combined with a detailed morphological analysis to seek characters that discriminate closely related species in the pleurocarpous moss genus, Rhytidiadelphus. & Duby) Gepp, P. caudatum Broth., P. quintasii Broth., P. molliculum Broth., P. madagassum Kiaer ex Besch., Porothamnium variifolioides J.L. The evolution of several gametophyte features previously thought to be reliable for delineating the family are also strongly correlated with habitat. M.Fleisch. Download. Traditionally, pleurocarpous mosses have been divided into three orders based mainly on their sporophytic characters. We identified Long H1, Short S1 and Long Y1 unit classes in nine Campeiostachys or Elymus species. plus Thaidium and some members of the Hypnaceae, (2) the members of the Plagiotheciaceae (in a relatively wide sense) and (3) the Hookeriales-Sematophyllaceae-Garovagliodeae. The alignment has 116 sequences and 2,738 aligned positions. De Sloover comb. A close relationship is suggested, phenetically at least if not cladistically, to the Thuidiaceae.